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Weight contributes to Horizontal Flight.-That the weight of the body plays an important part in the production of flight may be proved by a very simple experiment.

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If I take two primary feathers and fix them in an ordinary cork, as represented at fig. 55, and allow the apparatus to drop from a height, I find the cork does not fall vertically downwards, but downwards and forwards in a curve. follows, because the feathers a, b are twisted flexible inclined planes, which arch in an upward direction. They are in fact true wings in the sense that an insect wing in one piece is a true wing. (Compare a, b, c of fig. 55, with g, g', s of fig. 82, p. 158.) When dragged downwards by the cork (c), which would, if left to itself, fall vertically, they have what is virtually a down stroke communicated to them. Under these circumstances a struggle ensues between the cork tending to fall vertically and the feathers tending to travel in an upward direction, and, as a consequence, the apparatus describes the curve d e f g before reaching the earth h, i. This is due to the action and reaction of the feathers and air upon each other, and to the influence which gravity exerts upon the cork. The forward travel of the cork and feathers, as compared with the space through which they fall, is very great. Thus, in some instances, I found they advanced as much as a yard and a half in a descent of three yards. Here, then, is

an example of flight produced by purely mechanical appliances. The winged seeds fly in precisely the same manner. The seeds of the plane-tree have, e.g. two wings which exactly resemble the wings employed for flying; thus they taper from the root towards the tip, and from the anterior margin towards the posterior margin, the margins being twisted and disposed in different planes to form true screws. This arrangement prevents the seed from falling rapidly or vertically, and if a breeze is blowing it is wafted to a considerable distance before it reaches the ground. Nature is uniform and consistent throughout. She employs the same principle, and very nearly the same means, for flying a heavy, solid seed which she employs for flying an insect, a bat, or a bird.

When artificial wings constructed on the plan of natural ones, with stiff roots, tapering semi-rigid anterior margins, and thin yielding posterior margins, are allowed to drop from a height, they describe double curves in falling, the roots of the wings reaching the ground first. This circumstance proves the greater buoying power of the tips of the wings as compared with the roots. I might refer to many other experiments made in this direction, but these are sufficient to show that weight, when acting upon wings, or, what is the same thing, upon elastic twisted inclined planes, must be regarded as an independent moving power. But for this circumstance flight would be at once the most awkward and laborious form of locomotion, whereas in reality it is incomparably the easiest and most graceful. The power which rapidly vibrating wings have in sustaining a body which tends to fall vertically downwards, is much greater than one would naturally imagine, from the fact that the body, which is always beginning to fall, is never permitted actually to do So. Thus, when it has fallen sufficiently far to assist in elevating the wings, it is at once elevated by the vigorous descent of those organs. The body consequently never acquires the downward momentum which it would do if permitted to fall through a considerable space uninterruptedly. It is easy to restrain even a heavy body when beginning to fall, while it is next to impossible to check its progress when

it is once fairly launched in space and travelling rapidly in a downward direction.

Weight, Momentum, and Power, to a certain extent, synonymous in Flight.-When a bird rises it has little or no momentum, so that if it comes in contact with a solid resisting surface it does not injure itself. When, however, it has acquired all the momentum of which it is capable, and is in full and rapid flight, such contact results in destruction. My friend Mr. A. D. Bartlett informed me of an instance where a wild duck terminated its career by coming violently in contact with one of the glasses of the Eddystone Lighthouse. The glass, which was fully an inch in thickness, was completely smashed. Advantage is taken of this circumstance in killing sea-birds, a bait being placed on a board and set afloat with a view to breaking the neck of the bird when it stoops to seize the carrion. The additional power due to momentum in heavy bodies in motion is well illustrated in the start and progress of steamboats. In these the slip, as it is technically called, decreases as the speed of the vessel increases; the strength of a man, if applied by a hawser attached to the stern of a moderate-sized vessel, being sufficient to retard, and, in some instances prevent, its starting. In such a case the power of the engine is almost entirely devoted to "slip" or in giving motion to the fluid in which the screw or paddle is immersed. It is consequently not the power residing in the paddle or screw which is cumulative, but the momentum inhering in the mass. In the bird, the momentum, alias weight, is made to act upon the inclined planes formed by the wings, these adroitly converting it into sustaining and propelling power. It is to this circumstance, more than any other, that the prolonged flight of birds is mainly due, the inertia or dead weight of the trunk aiding and abetting the action of the wings, and so relieving the excess of exertion which would necessarily devolve on the bird. It is thus that the power which in living structures resides in the mass is conserved, and the mass itself turned to account. But for this reciprocity, no bird could retain its position in the air for more than a few minutes at a time. This is proved by the comparatively brief upward flight of the lark and the hovering of the hawk

when hunting. In both these cases the body is exclusively sustained by the action of the wings, the weight of the trunk taking no part in it; in other words, the. weight of the body does not contribute to flight by adding its momentum and the impulse which momentum begets. In the flight of the albatross, on the other hand, the momentum acquired by the moving mass does the principal portion of the work, the wings for the most part being simply rotated on and off the wind to supply the proper angles necessary for the inertia or mass to operate upon. It appears to me that in this blending of active and passive power the mystery of flight is concealed, and that no arrangement will succeed in producing flight artificially which does not recognise and apply the principle here pointed out.

Air-cells in Insects and Birds not necessary to Flight.-The boasted levity of insects, bats, and birds, concerning which so much has been written by authors in their attempts to explain flight, is delusive in the highest degree.

Insects, bats, and birds are as heavy, bulk for bulk, as most other living creatures, and flight can be performed perfectly by animals which have neither air-sacs nor hollow bones; airsacs being found in animals never designed to fly. Those who subscribe to the heated-air theory are of opinion that the air contained in the cavities of insects and birds is so much lighter than the surrounding atmosphere, that it must of necessity contribute materially to flight. I may mention, however, that the quantity of air imprisoned is, to begin with, so infinitesimally small, and the difference in weight which it experiences by increase of temperature so inappreciable, that it ought not to be taken into account by any one endeavouring to solve the difficult and important problem of flight. The Montgolfier or fire-balloons were constructed on the heated-air principle; but as these have no analogue in nature, and are apparently incapable of improvement, they are mentioned here rather to expose what I regard a false theory than as tending to elucidate the true principles of flight.

When we have said that cylinders and hollow chambers increase the area of the insect and bird, and that an insect

and bird so constructed is stronger, weight for weight, than one composed of solid matter, we may dismiss the subject; flight being, as I shall endeavour to show by-and-by, not so much a question of levity as one of weight and power intelligently directed, upon properly constructed flying surfaces.

The bodies of insects, bats, and birds are constructed on strictly mechanical principles,-lightness, strength, and durability of frame being combined with power, rapidity, and precision of action. The cylindrical method of construction is in them carried to an extreme, the bodies and legs of insects displaying numerous unoccupied spaces, while the muscles and solid parts are tunnelled by innumerable airtubes, which communicate with the surrounding medium by a series of apertures termed spiracles.

A somewhat similar disposition of parts is met with in birds, these being in many cases furnished not only with hollow bones, but also (especially the aquatic ones) with a liberal supply of air-sacs. They are likewise provided with a dense covering of feathers or down, which adds greatly to their bulk without materially increasing their weight. Their bodies, moreover, in not a few instances, particularly in birds of prey, are more or less flattened. The air-sacs are well seen in the swan, goose, and duck; and I have on several occasions minutely examined them with a view to determine their extent and function. In two of the specimens which I injected, the material employed had found its way not only into those usually described, but also into others which ramify in the substance of the muscles, particularly the pectorals. No satisfactory explanation of the purpose served by these air-sacs has, I regret to say, been yet tendered. According to Sappey,1 who has devoted a large share of attention to the subject, they consist of a membrane which is neither serous nor mucous, but partly the one and partly the other; and as blood-vessels in considerable numbers, as my preparations

1 Sappey enumerates fifteen air-sacs,-the thoracic, situated at the lower part of the neck, behind the sternum; two cervical, which run the whole length of the neck to the head, which they supply with air; two pairs of anterior, and two pairs of posterior diaphragmatic; and two pairs of abdo· minal.

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